1. Wahlberg N., Peña C., Ahola M., Wheat C. W., Rota J. 2016 PCR primers for 30 novel gene regions in the nuclear genomes of Lepidoptera. ZooKeys 596: 129–141. doi:10.3897/zookeys.596.8399
  2. Rota, J., Peña, C. and Miller, S. E. 2016. The importance of long-distance dispersal and establishment events in small insects: historical biogeography of metalmark moths (Lepidoptera, Choreutidae). Journal of Biogeography 43: 1254–1265. doi:10.1111/jbi.12721
  3. Papakostas S., Michaloudi E., Proios K., Brehm M., Verhage L., Rota J., Peña C., Stamou G., Pritchard V.L., Fontaneto D., Declerck S.A.J. 2016. Integrative taxonomy recognizes evolutionary units despite widespread mitonuclear discordance: evidence from a rotifer cryptic species complex. Systematic Biology 65(3): 508–524. doi:10.1093/sysbio/syw016
  4. Vesterinen, E. J., Ruokolainen, L., Wahlberg, N., Peña, C., Roslin, T., Laine, V. N., Vasko, V., Sääksjärvi, I. E., Norrdahl, K. and Lilley, T. M. 2016. What you need is what you eat? Prey selection by the bat Myotis daubentonii. Molecular Ecology 25: 1581–1594. doi:10.1111/mec.13564
  5. Peña, C., Witthauer, H., Klečková, I., Fric, Z. & Wahlberg, N. 2015. Adaptive Radiations in Butterflies: Evolutionary History of the Genus Erebia (Nymphalidae: Satyrinae). Biological Journal of the Linnean Society, 116(2): 449–467. doi:10.1111/bij.12597
  6. Peña, C. & Espeland, M. 2015. Diversity dynamics in Nymphalidae butterflies: Effect of phylogenetic uncertainty on diversification rate shift estimates. PLoS ONE, 10(4): e0120928. doi:10.1371/journal.pone.012092
  7. Matos-Maraví, P. F., Núñez Águila, R., Peña, C., Miller, J. Y., Sourakov, A. & Wahlberg, N. 2014. Causes of endemic radiation in the Caribbean: evidence from the historical biogeography and diversification of the butterfly genus Calisto (Nymphalidae: Satyrinae: Satyrini). BMC Evolutionary Biology, 14: 199. doi:10.1186/s12862-014-0199-7.
  8. PDF Gómez, I. C., Sääksjärvi, I. E., Broad, G. R., Puhakka, L., Castillo, C., Peña C. & Pádua, D. G. 2014. The Neotropical species of Xanthopimpla Saussure (Hymenoptera: Ichneumonidae: Pimplinae). Zootaxa, 3774(1): 57–73. doi:10.11646/zootaxa.3774.1.4
  9. Wahlberg, N., Wheat, C. W. & Peña, C. 2013: Timing and patterns in the taxonomic diversification of Lepidoptera (butterflies and moths). PLoS ONE, 8(11): e80875. doi:10.1371/journal.pone.0080875
  10. PDF Matos-Maravi, P. F., Peña, C., Willmott, K. R., Freitas, A. V. L. & Wahlberg, N. 2013. Systematics and evolutionary history of butterflies in the "Taygetis clade" (Nymphalidae: Satyrinae: Euptychiina): towards a better understanding of Neotropical biogeography. Molecular Phylogenetics and Evolution, 66(1): 54–68. doi:10.1016/j.ympev.2012.09.005
  11. PDF Peña, C. & Malm, T. 2012. VoSeq: a Voucher and DNA Sequence Web Application. PLOS ONE, 7(6): e39071. | doi:10.1371/journal.pone.0039071
  12. PDF Nyman, T., Linder, H. P., Peña, C., Malm, T. & Wahlberg, N. 2012. Climate-driven diversity dynamics in plants and plant-feeding insects. Ecology Letters, 15: 889–898. | doi:10.1111/j.1461-0248.2012.01782.x
  13. PDF Heikkilä, M., Kaila, L., Mutanen, M., Peña, C., & Wahlberg, N. 2012. Cretaceous origin and repeated Tertiary diversification of the redefined butterflies. Proceedings of the Royal Society B, 279: 1093–1099. | doi:10.1098/rspb.2011.1430
  14. PDF Peña, C. 2011. Métodos de inferencia filogenética. Revista peruana de Biología, 18(2): 265–267. [Commentary]
  15. PDF Pulido-B, H. W., Andrade-C, M. G., Peña, C., & Lamas, G. 2011. Two new taxa of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the Andes of Colombia and Peru. Zootaxa, 2906: 43–51.
  16. PDF Marín, M. A., Peña, C., Freitas, A. V. L., Wahlberg, N. & Uribe, S. I. 2011. From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera, Nymphalidae): history, progress and prospects. Neotropical Entomology, 40(1):1–13. | doi:10.1590/S1519-566X2011000100001
  17. PDF Peña, C., Nylin, S., & Wahlberg, N. 2011. The radiation of Satyrini butterflies (Nymphalidae: Satyrinae): a challenge for phylogenetic methods. Zoological Journal of the Linnean Society, 161(1): 64–87. | doi: 10.1111/j.1096-3642.2009.00627.x
  18. PDF Peña, C., Nylin, S., Freitas, A.V.L. & N. Wahlberg. 2010. Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta, 39: 243–258. | doi:10.1111/j.1463-6409.2010.00421.x
  19. PDF Kodandaramaiah, U., Peña, C., Braby, M., Grund, R., Müller, C. J., Nylin, S. & N. Wahlberg. 2010. Phylogenetics of Coenonymphina (Nymphalidae: Satyrinae) and the problem of rooting rapid radiations. Molecular Phylogenetics and Evolution, 54: 386–394. | doi:10.1016/j.ympev.2009.08.012
  20. PDF Wahlberg, N., Leneveu, J., Kodandaramaiah, U., Peña, C., Nylin, S., Freitas, A. V. L. & Brower, A. V. Z. 2009. Nymphalid butterflies diversify following near demise at the Cretaceous/Tertiary boundary. Proceedings of the Royal Society Series B, 276: 4295–4302. | doi:10.1098/rspb.2009.1303. Electronic Material, download here. Read the interview in ScienceNOW.
  21. PDF Peña, C., & N. Wahlberg. 2008. Prehistorical climate change increased diversification of a group of butterflies. Biology Letters 4(3): 274–278. | Supplementary Material | Commentary in Telegraph | doi:10.1098/rsbl.2008.0062 |
  22. PDF Wahlberg, N. & C. Peña. 2007. A Dominican Republic sojourn. News of the Lepidopterists' Society 49(1): 12–15, 37.
  23. PDF Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, A.V.L. Freitas & A.V.Z. Brower. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution 40(1): 29–49. | dataset in TNT format | doi:10.1016/j.ympev.2006.02.007 |
  24. PDF Freitas, A.V.L. & C. Peña. 2006. Description of genus Guaianaza for "Euptychia" pronophila (Lepidoptera: Nymphalidae: Satyrinae) with a description of the immature stages. Zootaxa 1163: 49–59.
  25. PDF Peña, C. & G. Lamas. 2005. Revision of the butterfly genus Forsterinaria Gray, 1973 (Lepidoptera: Nymphalidae, Satyrinae). Revista peruana de Biología 12(1): 5–48.


PDF Peña, C. 2009. Evolutionary history of the butterfly subfamily Satyrinae (Lepidoptera: Nymphalidae). Department of Zoology, Stockholm University, Stockholm, SWEDEN. March 2009. PhD thesis. ISBN: 9789171558107

PDF Peña, C. 2007. Butterflies and grasses: evolutionary history of the subfamily Satyrinae. Department of Zoology, Stockholm University, Stockholm, SWEDEN. April 2007. Licentiate thesis.

PDF Peña, C. 2004. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data, A preliminary study. Department of Zoology, Stockholm University, Stockholm, SWEDEN. October 2004. Examensarbete thesis.

Peña, C. 2004. "Systematics of the genus Forsterinaria Gray, 1973 (Lepidoptera: Nymphalidae, Satyrinae)" [in Spanish; Abstract in English]. Diploma in Biology, Major Zoology, Department of Zoology, Faculty of Biological Sciences, Universidad Nacional Mayor de San Marcos, Lima, PERU, April 2004. Thesis for degree of Biologist.